Etymology
The word "stipule" comes from the Latin "stipula," which means "stalk" or "straw."
Meaning
In botany, a stipule is a pair of modified leaves or leaf-like structures that are located at the base of a petiole (leaf stalk). Stipules can vary greatly in shape and size, from small and inconspicuous to large and showy.
Origin
Stipules are thought to have evolved from the stipules of the first terrestrial plants, which were likely non-vascular plants such as mosses and liverworts. In these plants, stipules helped to protect the young and delicate leaves from damage. As plants evolved and became more complex, stipules took on additional functions, such as photosynthesis, water storage, and defense against herbivores.
Significance
Stipules can be an important taxonomic identifier for certain species of plants. The presence, absence, shape, and size of stipules are often used to distinguish one species from another. Additionally, stipules play various roles in the growth and development of plants. They can help to support leaves and stems, protect young leaves, and regulate water and nutrient uptake.
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